TAC found http://tools.cisco.com/Support/BugToolKit/search/getBugDetails.do?method=fetchBugDetails&bugId=CSCsm15661
Problem Description:
When upgrading from 4.x to 5.x, settings for Standard and Closed Transfer rules for (non-Subscriber) Call Handlers may be overwritten with the settings from the Alternate Transfer rule. This may result in call transfer rules that were set to "Yes, ring this extension" before the upgrade to "No (Send Directly to Greeting) after the upgrade.
Workaround
a) Set the extension manually
b) Use the Bulk Edit tool
The fix is not available yet for 5.x or 7.x