When you choose the option "Enable Signature and Engine Updates from Cisco.com", the Cisco.com URL will automatically be filled in with the correct URL. Unless you don't have it, it's "https://198.133.219.25//cgi-bin/front.x/ida/locator/locator.pl".
The username and password would be your cisco CCO account.
It will use the IPS management ip address to connect to cisco.com, so the IPS mgmt ip address needs to have Internet connectivity.
Hope that helps.
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